Rescuing a squirrel after an accident involving a vacuum cleaner, comic-reading cynic Flora Belle Buckman is astonished when the squirrel, Ulysses, demonstrates astonishing powers of strength and flight after being revived.

Two warrior squirrels lead the battle against Gulo, a bloodthirsty wolverine who will stop at nothing to recover the Walking Stone that will give him the authority to rule the lands of ice beyond the Great Sea.